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A particularly critical driver of cell ageing in various lower species such as Caenorhabditis elegans 27,28 and probably mammals 29 is declining lysosomal function linked to failing intralumenal acidification 26 , which is believed to be responsible for the initial build-up of multiple pathogenic aggregation-prone proteins (β-amyloid precursor protein (APP) peptides (for example, amyloid-β and APP carboxy-terminal fragment (APP-βCTF), TAR DNA-binding protein 43 (TDP43), synuclein)) in the brain upon normal ageing

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Sekine K, Nagata N, Sakamoto K, Arai T, Shimbo T, Shinozaki M,